In addition to cleaning out the last leftovers from SeaMonkey, the most noticeable and useful change is the fact that eee-control now is accessible from a quick launch icon on the task bar. One stop shopping for most eee-specific settings, including over-clocking if you like to burn your battery faster.
Other useful additions include a battery meter and CPU temperature gauge on the task bar... almost makes my install of pwidgets redundant, but the eye candy is soo pretty...
Overall the most important improvement of PupEee 4.32 over previous versions is the change to Chrome and pwireless2 - those two changes alone is worth the trouble of upgrading.
PupEee 4.31 is discussed in this thread at EeeUser.com, has it's own forum here, and can be downloaded from this site.
Above: Beta 5, with eee-control, powertop and chrome running
Below: The various functions on the task bar